Denis Leary Diego (voice)
John Leguizamo Sid (voice)
Queen Latifah Ellie (voice)
Alan Tudyk Cholly (voice)
Will Arnett Lone Gunslinger Vulture (voice)
Stephen Root Aardvark Dad (voice)
Seann William Scott Crash (voice)
Jon Vitti Story Consultant
Josh Peck Eddie (voice)
Ariel Winter Additional Voices (voice)
Jay Leno Fast Tony (voice)
Carlos Saldanha Director, Dodo (voice)
Ray Romano Manny (voice)
Bill Farmer Additional Voices (voice)
Debi Derryberry Diatryma Mom (voice)
Jack Angel Additional Voices (voice)
Bob Bergen Additional Voices (voice)
Mindy Sterling Female Ox (voice)
Jess Harnell Additional Voices (voice)
Chris Renaud Story Artist
Joseph Bologna Mr. Start (voice)
John Powell Original Music Composer
Renée Taylor Mrs. Start (voice)
Laura Marano Additional Voices (voice)
Paul Butcher Additional Voices (voice)
Rodger Bumpass Additional Voices (voice)
Jansen Panettiere Shovelmouth Boy (voice)
Danny Mann Additional Voices (voice)
Chris Wedge Scrat (voice), Executive Producer
Randy Thom Sound Designer, Supervising Sound Editor, Sound Mixer
Chris Meledandri Executive Producer
Bob Camp Story Artist
Chris Scarabosio Sound Mixer
Clea Lewis Female Mini Sloth (voice)
Madeleine Martin Additional Voices (voice)
James Sie Freaky Male (voice)
Laraine Newman Additional Voices (voice)
Jennifer Darling Additional Voices (voice)
Peter Ackerman Dung Beetle Dad (voice)
David Acord Assistant Sound Editor
Al Nelson Sound Effects Editor
Michael Thurmeier Visual Effects, Supervising Animator
Brandon Proctor Sound Mix Technician
Shawn Murphy Scoring Mixer
Jim Hecht Screenplay
Mickie McGowan Additional Voices (voice)
Nick Bruno Animation
Gwendolyn Yates Whittle Supervising ADR Editor, Supervising Sound Editor
Lee Ryan Songs
Will Files Assistant Sound Designer
John Cygan Additional Voices (voice)
Lora Hirschberg Sound Mixer
Brett Hoffman Script Coordinator, Story Coordinator
Gerry Swallow Screenplay, Story
Harrison Fahn Additional Voices (voice)
Peter Gaulke Story, Screenplay
Zack Shada Additional Voices (voice)
Crispin Freeman Additional Voices (voice)
Chris Gilligan Story Artist
Jan Rabson Additional Voices (voice)
Marshall Efron Start Dad (voice)
Sean England Foley Recordist
Peter de Sève Condor Chick (voice), Character Designer
Jim Passon Color Timer
Gary Burritt Negative Cutter
Ellen Heuer Foley Artist
Gregory Romano Start Boy 1 (voice)
Raul Chavez Animation
Olivia DeLaurentis Additional Voices (voice)
Shane Baumel Additional Voices (voice)
Matt Romano Start Boy 2 (voice)
Craig Paulsen Associate Editor